April 2026 – present: Postdoctoral Scholar – Kenchanmane-Raju Lab, UC Riverside

July 2023 – Mar 2026: Postdoctoral Researcher – Jackson Lab, Cold Spring Harbor Laboratory

Nov 2017 – July 2023: Doctoral Student – Schnurbusch Lab, IPK Gatersleben

Publications
1. Iohannes, S. D., Shanmugaraj, N., & Jackson, D. ‘‘Quantitative imaging of RNA in plants using hybridization chain reaction-RNA fluorescence in situ hybridization (HCR RNA FISH)’’. Current Protocols, 2026.

2. Rutten, T., Thirulogachandar, V., Huang, Y., Shanmugaraj, N., Koppolu, R., Ortleb, S., Hensel, G., Kumlehn, J., Melzer, M., Schnurbusch, T., ‘‘Anatomical insights into the vascular layout of the barley rachis: implications for transport and spikelet connection’’, Annals of Botany, 133(7), 2024, Pages 983– 996.

3. Shanmugaraj, N., Rajaraman, J., Sandip, K., Kamal, R., Huang, Y., Thirulogachandar, V., Budhagatapalli, N., Moya, Y. A.T., Hajirezaei, M.R., Garibay, A., Rutten, T., Kumlehn, J., Hensel, G., Melzer, M., von Wirén, N., Mock, H.P., and Schnurbusch, T., Multilayered regulation of developmentally programmed pre-anthesis tip degeneration of the barley inflorescence". The Plant Cell. 35(11), 2023, Pages 3973–4001.

4. Shanmugaraj, N., Svatos, A., Twan, R., Mock, H.P., and Schnurbusch, T., Fast and reproducible matrix deposition for MALDI mass spectrometry imaging with improved glass sublimation setup, Journal of the American Society for Mass Spectrometry, 2023 34(3): 513-517.

5. Huang, Y., Kamal, R., Shanmugaraj, N., Rutten, T., Thirulogachandar, V., Zhao, S., Hoffie, I.,Hensel, G., Rajaraman, J., Moya1, Y. A. T., Hajirezaei, M-R., Himmelbach, A., Poursarebani, N.,Lundqvist, U., Kumlehn, J., Stein, N., von Wirén, N., Mascher, M., Melzer, M., Schnurbusch, T., A molecular framework for grain number determination in barley,. Science Advances,