Nearly every aspect of growth, morphogenesis, physiology, and stress response is influenced by cell/tissue-type specific gene expression. Transcription factors (TFs) recognize cis-regulatory elements and signal transcription machinery for gene regulation, and the interaction between TFs and their target genes creates spatiotemporal patterns of gene expression. Accurate functional annotation of these regulatory sites is essential for a comprehensive understanding of gene regulation in a variety of cell types and under diverse conditions…